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Lebanese amber

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Tragichrysa ovoruptora, a green lacewing larvae

Lebanese amber is fossilized resin found in Lebanon and surrounding countries. A majority of the amber with fossil inclusions has been found within 3 layers of the Grès du Liban alloformation, which dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. Upper Jurassic Kimmeridgian amber has also been found in Lebanon from the Bhannes formation, but so far only one locality, Aintourine, has produced inclusions.[1] The main amber deposit was formed on what was then the northern coast of Gondwana, probably within a tropical or subtropical zone in a temperate or hot climate.[2] Analysis of the sediment from the site indicate the amber was produced in coastal, possibly estuarine conditions.[1]

It is the oldest source of amber with a significant number of inclusions. Up to 300 sources of Lebanese amber have been recovered and 17 of them are important sources of organic inclusions, which are the oldest of their kind. The inclusions help to document Cretaceous fauna and flora.

Tyruschrysa melqart, a green lacewing larvae

Origins

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Lebanese amber can be found in Lebanon and neighboring areas of the Levant. Up to 300 different sources of amber had been discovered by 2010. The amber was deposited in the Cretaceous era and is rich in fossil synclusions. 19 of the discovered sources are rich in inclusions from the Early Cretaceous. All of them are located in Lebanon, which makes it the largest source of inclusion from that period.[3]

History

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Aside from possible early reports of Phoenician usage, the oldest reports of Lebanese amber are from 19th-century accounts, these tended only to be incidental due to Lebanese amber's gemological quality in comparison to Baltic amber, and the local people were more interested in the associated lignite as a source of fuel.[4]

Properties

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Lebanese amber can be found in a vast variety of colors such as yellow, orange, dark red or iridescent jet black. Rarely in white, milky or cream. The variation of color tone is caused by the air contained in the amber. The density of Lebanese amber is 1.054 g/cm3. It tends to be fragile and easy to damage.[5]

Inclusions

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Inclusions are quite common for Lebanese amber, which is rich in organisms of the Barremian epoch. Next to Jordanian amber, Lebanese amber is the oldest amber to have yielded significant invertebrate inclusions alongside the Wealden amber from the equivalently aged Wessex Formation of the UK, which is much less productive. Organisms preserved in Lebanese amber are dated back to the period prior to the angiosperm radiation, which was the period of massive extinction of old groups of arthropods, as well as the emergence of the new ones, some of which co-evolved with angiosperms. The organisms are preserved in good condition and shape.[4] The diversity and number of co-inclusions help to draw conclusions about mutual relations and co-existence.[6]

Paleobiota

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After Poinar & Milki (2001),[2] Maksoud & Azar (2020)[7] and subsequent studies.

Bacteria

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Genus Species Notes Images

Streptomyces

sp.

Fungi

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Genus Species Notes Images

Chytridiales

indet.

Mucorales

indet.

Plants

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Genus Species Notes Images

Agathis

levantensis An araucarian tree responsible for the production of amber.

Hexapoda

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Genus Species Notes Images

Archaeatropos

An archaeatropid psocodean.[8]

Arthropleona

indet. A springtail.

Asphaeropsocites

A sphaeropsocid psocodean.[9]

Aphelopus

A dryinid wasp.

Archiaustroconops

A ceratopogonid fly.

Archiculicoides

A ceratopogonid fly.

Archisciada

A sciadocerid fly.

Atelestites

An empidid fly.

Austroconops

A ceratopogonid fly.

Avenaphora

A dolichopodid fly.

Banoberotha

A beaded lacewing.

Bcharreglaris

An archaeatropid psocodean.[8]

Bernaea

A whitefly.

Blattodea

indet. A cockroach.[9]

Chomeromyia

A fly.

Conovirilus

A leptophlebiid mayfly.

Corethrella

A corethrellid biting midge.

Cretaceomachilis

A meinertellid bristletail.

Cretacetrocta

A pachytroctid psocodean.[8]

Cretapsychoda

A psychodid fly.

Enicocephalinus

An enicocephalid.

Exitelothrips

A scudderothripid thrips.

Fossileptoconops

A ceratopogonid fly.

Glaesoconis

A dustywing.

Heidea

A whitefly.

Incurvariites

A incurvariid moth.
Jankotejacoccus A jankotejacoccid scale insect.[10]

Jezzinothrips

A jezzinothripid thrips.

Lebambromyia

A phorid fly.

Lebanaphis

A tajmyraphidid hemipteran.

Lebania

A tipulid crane fly.

Lebanoconops

A ceratopogonid fly.

Lebanoculicoides

A ceratopogonid fly.

Leptoconops

A ceratopogonid fly.

Libaneuphoris

A libanophorid pscocodean.

Libanobythus

A scolobythid hemipteran.

Libanochlites

A non-biting midge.

Libanoculex

A chaoborid fly, previously interpreted as a mosquito.[11]

Libanoglaris

An archaeatropid psocodean.[8]

Libanophlebotomus

A psychodid fly.

Libanomphientomum

A electrentomoid psocodean.[8]

Libanopsychoda

A psychodid fly.

Libanopsyllipsocus

A psyllipsocid psocodean.[8]

Libanorhinus

A nemonychid weevil.

Libanosemidalis

A dustywing.

Lonchopterites

A lonchopterid fly.

Lonchopteromorpha

A lonchopterid fly.

Rhadinolabis

An earwig.[9]

Setoglaris

An archaeatropid psocodean.[8]

Lebanotermes

A termite.[9]

Megarostrum

A tajmyraphidid hemipteran.

Mesobolbomyia

A rhagionid fly.

Mundopoides

A cixiid.

Neocomothrips

A neocomothripid thrips.

Paleochrysopilus

A rhagionid fly.

Paleopsychoda

A psychodid fly.

Palaeosiamoglaris

A prionoglarid psocodean.[8]

Parasabatinca

A micropterigid moth.

Paraberotha

A beaded lacewing.

Paramesopsocus

A electrentomid psocodean.[8]

Phaetempis

An empidid fly.

Phlebotomites

A phlebotomid fly.

Progonothrips

A rhetinothripid thrips.

Protoculicoides

A ceratopogonid fly.

Protopsychoda

A psychodid fly.

Psocodea

sp. 1 A juvenile psocodean.[8]

Raptorapax

A rhachiberotid neuropteran.[9]

Rhetinothrips

A rhetinothripid thrip.

Scaphothrips

A scaphothripid thrips.

Scudderothrips

A scudderothripid thrips.

Sphaeropsocites

A sphaeropsocid psocodean.[8]

Trichinites

An empidid fly.

Yuripopovina

A yuripopovinid hemipteran.[9]

Arachnida

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Genus Species Notes Images

Acari

indet. Various free-living mites.

Anystidae

sp. An anystid mite.

Archaeobuthus

An archaeobuthid scorpion.[9]

Erythraeoidea

indet. A erythraeid mite.

Erythraeoidea

indet. Leptus sp.?

Lebansegestria

A segestriid spider.

Lebanoecobius

A oecobiid spider.

Linyphiidae

sp. A linyphiid spider.

Microsegestria

A segestriid spider.

Neoliodes

A neoliodid mite.

Oonopidae

sp. An oonopid spider.

Palaeomicromenneus

A deinopid spider.

Plumorsolus

A plumorsolid spider.

Pseudoscorpiones

indet. A pseudoscorpion.

Zamilia

A oecobiid spider.

Other invertebrates

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Genus Species Notes Images

Chilopoda indet.

Cretaciomermis

A nematode.

Electroxenus

A polyxenid millipede.

Heleidomermis

A mermithid nematode.

Libanoxenus

A polyxenid millipede.

Pupilloidea indet.

A possible pupillid snail.

Vertebrates

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Vertebrates of the Lebanese Amber
Genus Species Notes Images

Aves[12]

indeterminate A single feather and the only known bird remains of the biota.

Baabdasaurus[13]

B. xenurus Autarchoglossan lizard.

References

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  1. ^ a b Maksoud, Sibelle; Tabakian, Haig (2025-12-01). "Lebanese amber: A time capsule from the dawn of modern ecosystems". Palaeoworld. 34 (6) 201015. doi:10.1016/j.palwor.2025.201015. ISSN 1871-174X.
  2. ^ a b Raif Milki, George Poinar, Lebanese Amber: The Oldest Insect Ecosystem in Fossilized Resin, 2001. ISBN 978-0-87071-533-4
  3. ^ David I. Green, David Penney, Fossils in Amber: Remarkable Snapshots of Prehistoric Forest Life, Siri Scientific Press, 2011
  4. ^ a b Penney, David (2010). "Chapter 14: Lebanese Amber". Biodiversity of fossils in amber from the major world deposits. Siri Scientific Press. ISBN 978-0-9558636-4-6. OCLC 904772774.
  5. ^ Andrew Ross, Amber, Harvard University Press, 1998
  6. ^ George Poinar, Life in amber, Stanford University Press, 1992
  7. ^ Maksoud, Sibelle; Azar, Dany (2020-04-30). "Lebanese amber: latest updates". Palaeoentomology. 3 (2): 125–155. Bibcode:2020Plegy...3..125M. doi:10.11646/palaeoentomology.3.2.2. ISSN 2624-2834.
  8. ^ a b c d e f g h i j k Marina, Hakim; Dany, Azar (2024). "Contributions to the Palaeobiodiversity of Psocodea ('Psocoptera') from Lebanese Amber: A Review". Fossil Studies. 2 (3). doi:10.3390/fo (inactive 1 July 2025). ISSN 2813-6284. Archived from the original on 2024-12-03. Retrieved 2025-06-15.{{cite journal}}: CS1 maint: DOI inactive as of July 2025 (link)
  9. ^ a b c d e f g Azar, D. (2012). Lebanese amber: a "Guinness Book of Records". Annales Universitatis Paedagogicae Cracoviensis. Studia ad Didacticam Biologiae Pertinentia, 2(I), 44-60.
  10. ^ Peter, Vršanský; Hemen, Sendi; Júlia, Kotulová; Jacek, Szwedo; Martina, Havelcová; Helena, Palková; Lucia, Vršanská; Jakub, Sakala; L'ubica, Puškelová; Marián, Golej; Adrian, Biroň; Daniel, Peyrot; Donald, Quicke; Didier, Néraudeau; Pavel, Uher (2025-02-07). "Jurassic Park approached: a coccid from Kimmeridgian cheirolepidiacean Aintourine Lebanese amber". National Science Review. 12 (3). doi:10.1093/n (inactive 29 September 2025). ISSN 2095-5138. Archived from the original on 2025-05-05. Retrieved 2025-07-12.{{cite journal}}: CS1 maint: DOI inactive as of September 2025 (link)
  11. ^ Harbach, Ralph E. (2024-03-12). "Libanoculex intermedius is not a mosquito (Diptera: Culicidae): It is a chaoborid (Chaoboridae)". Zootaxa. 5424 (1): 139–144. doi:10.11646/zootaxa.5424.1.9. ISSN 1175-5334. PMID 38480294. Archived from the original on 9 August 2024. Retrieved 9 August 2024.
  12. ^ Schlee, D (1973). "Harzkonservierte fossile Vogelfedern aus der untersten Kreide [Resin-preserved fossil bird feathers from the lowermost Cretaceous]". Journal für Ornithologie. 114 (2): 207-219. doi:10.1007/BF01641171.
  13. ^ Arnold, E. N.; Azar, D.; Ineich, I.; Nel, A. (September 2002). "The oldest reptile in amber: a 120 million year old lizard from Lebanon". Journal of Zoology. 258 (1): 7–10. doi:10.1017/S0952836902001152.